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A new genus and species of Stenomylinae (Camelidae, Artiodactyla) from the Moonstone Formation (late Barstovian–early Hemphillian) of central Wyoming

Department of Geology and Geophysics, The University of Wyoming, Laramie, Wyoming 82071, U.S.A.

email: mcassil{at}uwyo.edu

Wyomylus whitei is a new genus and species of stenomyline camel from the Moonstone Formation (late Barstovian–earliest Hemphillian, middle–late Miocene) of central Wyoming. W. whitei, the youngest known species of the Stenomylinae, is a small, gracile camel that is primitive in many of its characters for such a late-appearing representative. Among the diagnostic characters of W. whitei are: extreme reduction of P1–P3; reduction of P4–M1; reduction of p4–m1; small posterior heel on M3; extreme nasal retraction; large and very deep, but unpocketed anterior maxillary fossa; large, very deep and slightly pocketed preorbital fossa; anteromaxillary and preorbital fossae confluent via an elongate depression directly ventral to the premaxilla–maxilla suture; posteriorly placed orbits; small, but well-developed wing-shaped internal pterygoid processes; very narrow dorsal moiety of the supraoccipital; very weak lambdoidal crest that does not overhang the occiput; and partially fused metatarsals III and IV.

Key Words: Barstovian • Camelidae • Clarendonian • Hemphillian • internal pterygoid processes • maxillary fossae • metapodials • Miocene • Moonstone Formation • Stenomylinae • Wyoming • Wyomylus whitei